OMAHA, Neb. — The remains of a sailor from Nebraska killed in the 1941 attack on Pearl Harbor have been identified.

The Defense POW/MIA Accounting Agency said Friday in a news release that the remains of Navy Storekeeper 2nd Class Gerald Clayton, of Central City, had been accounted for on Sept. 10.

Clayton was 21 and assigned to the battleship Oklahoma when the ship was attacked by Japanese planes on Dec. 7, 1941. The attack on the ship resulted in the deaths of 429 crewmen, including Clayton.

Clayton was among the unidentified remains buried in 46 plots at the National Memorial Cemetery of the Pacific, known as the Punchbowl, in Honolulu.

In 2015, agency personnel began exhuming the remains for analysis, using new DNA technology, to identify those lost.